Tweddle, D S C, Willoughby, N G, J.L.B. Smith Institute of Ichthyology. J.L.B. Smith Institute of Ichthyology; 1982. The distribution and identification of mormyrid fishes in Malawi, with notes on the synonymy of Marcusenius nyasensis and M. livingstonii (Mormyriformes: Mormyridae).

Tweddle, D S C, Lewis, Digby S C, Willoughby, N G. J.L.B. Smith Institute of Ichthyology, Rhodes University, Grahamstown, South Africa; 1979. The nature of the barriers separating the Lake Malawi and Zambezi fish faunas.